The escalation comes as Syria's six-year-old war becomes ever more complex, with US forces and their allies converging on the northern IS bastion of Raqa in close proximity to Russian-backed regime troops.
Further complicating matters, Iran said it launched missile strikes Sunday against "terror bases" in Syria's northeastern Deir Ezzor province in revenge for deadly attacks on its capital claimed by IS.
The US-backed Syrian Democratic Forces -- an alliance of Kurdish and Arab fighters -- is battling to oust the jihadists from Raqa, and broke into the IS stronghold city last week.
Government forces are not involved in the battle for Raqa, but they are advancing in an area southwest of the city, skirting around SDF fighters, their eyes set on the oil-rich province of Deir Ezzor.
"Aircraft from the 'international coalition' targeted one of our fighter planes in the Resafa region of southern Raqa province this afternoon while it was conducting a mission against the terrorist Islamic State group," said the army.
It warned of "the grave consequences of this flagrant aggression".
The US-led coalition later confirmed it shot down the Syrian warplane it said had dropped bombs near SDF forces.
"At 6:43 pm (1743 GMT), a Syrian regime SU-22 dropped bombs near SDF fighters... In accordance with rules of engagement and in collective self-defence of Coalition partnered forces, was immediately shot down by a US F/A-18E Super Hornet," the Combined Joint Task Force said in a statement.
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